从在火星所发现的神秘之箱－“潘朵拉之箱”被打开的瞬间，突然地出现巨大墙壁开始本作的故事。
被称之为“天空之壁”的墙壁，将日本列岛分割成三块，分别为“东都”、“西都”、“北都”及各自的首都也因此诞生了。 其中于“东都”上突然出现被命名为《SMASH》的未确认生命体....。
它们暗中地将人类逐渐逼进绝境。在那样的Smash 面前站出来一名青年－桐生战兔，其腰上系着腰带，他的手上握有着不可思议的瓶子。
“变身！” - 天才物理学家的青年变身成假面骑士Build，阻挡在Smash 的面前！
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