这是一个属于国王和王后的年代, 电影讲述了发生在美人鱼和路易十四国王之间一段独一无二的奇幻故事……法国, 路易十四建立起了强大的太阳帝国, 却只有一个低能儿的王子可以继承他的王位. 对于未来忧心忡忡的他, 经多方探访得知获得美人鱼的力量可以获得生命的永恒，确保王位的稳固。他派出得力干将伊夫斯,带领法国海军出海苦苦搜捕，终于抓得一只美人鱼,并带回凡尔赛宫软禁。与此同时, 美丽天真的女孩子玛丽.约瑟夫, 被人从海边修道院送入了皇宫。她其实是路易十四与最心爱的情妇的私生女,为了解决一笔战争欠款路易十四为她安排了一场联姻。初到皇室的玛丽对一切阴谋一无所知, 却邂逅了被禁皇宫的美人鱼并和她成为了好友，并逐渐爱上了看守美人鱼的军官伊夫斯。随着玛丽和美人鱼之间的关系越来越紧密, 玛丽决定反抗父亲, 冒着失去生命和自由的危险, 救出她的朋友…
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