人到中年，因为处在重要的心理和身体的转变期，事业与家庭的双重压力往往会让他们觉得喘不过气来，这个时候的人更能反思社会与自身，他们为碌碌无为的过去而痛心，更为那充满了无法预期的可能性的未来而焦虑:人就像天地间最渺小的一粒沙子，那种无助的感觉，如影随行。故事中所讲述的四个人，无一例外都在经历着中年危机:夹在神经质的老婆与比自己还要倨傲的儿子中间，道格感觉自己就要窒息了;而道格的高中好友鲍比也在经历着相似的生活，这个惧内的家庭妇男，被妻子像对待狗一样呼来喝去，男性的尊严被无数次践踏;达德利则是一个工作高度紧张的电脑操作员，也是四人中惟一未婚的男子，然而他被女性挫败无数次之后，已经丧失了对自己男性能力的信心，所有接近异性的计划都不可避免地在一场灾难性的混乱中画上了句点;伍迪看起来很富有，可是他的名模妻子正打算和他离婚，同时还要把他的全部家当做赡养费……
几年来，这四个好友每到周末都会穿上锃亮的皮衣，骑着摩托车跑上那么几圈，这一次，四个人一合计，决定彻底摆脱紧张的工作和家庭的责任压力，为自己放一周的假，骑着摩托车从辛辛那提出发，一直开到加州，他们带着足够的食物，就上路了。
他们以天为被、以地为床，枕着青草与泥土、看着星星入睡，想想都觉得充满着浪荡江湖的豪气……然而，这个搭配不是非常协调的四人组，离他们熟悉的环境越来越远、开始了他们生命中惟一的探险--征服公路，却遭遇了一个当初想都想不到的全新情况，使得这场旅行最终变成了挑战他们集体智慧与运气的一次变故。尤其是在他们碰到了真正的飞车党"Del Fuegos"后，被嘲弄的就不仅仅是他们生疏的车技了，还有他们的勇气与道德观，最终竟然阴差阳错地成就了他们的英雄梦，让达德利顺利告别单身。
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